Lion kills man at Johannesburg Zoo

“He had been with us for so long and had an amazing bond with the animals,” she said.

The lioness attacked Mr Ramonetha in a passageway which the creatures cannot normally access, suggesting a gate was left open, she added.

Afterwards, she was sedated with a tranquilliser dart and returned to her enclosure.

“There will be a full investigation and we will decide then what to do with Nyanga but it’s clear that this was just an instinctive reaction on her part, perhaps because he tried to run away. She didn’t take his body away like prey – she bit him once in the neck and that was it.

“It’s just an unfortunate accident and devastating for all of us.”

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